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Publications

Italics indicates graduate students, * plus italics indicate students for whom I am the primary advisor, †contact author/senior author, ^undergraduate students

2017

1. Smith, R.L.*, K.H. Beard, and A.B. Shiels. 2017. Different prey resources suggest little competition between non-native frogs and insectivorous birds despite isotopic niche overlap. Biological Invasions. 19(3):1001-1013. 
2. Kulmatiski, A., K.H. Beard, J. Norton, J. Heavilin, L. Forero, and J. Grenzer. 2017. Live long and prosper: plant-soil feedback, lifespan and landscape abundance covary. Ecology
3. Marsh, D.M., B.J. Cosentino, K.S. Jones, J.J. Apodaca, K.H. Beard, J.M. Bell, C. Bozarth, D. Carper, J.F. Charbonnier, A. Dantas, E.A. Forys, M. Foster, J. General, K.S. Genet, M. Hanneken, K.R. Hess, S.A. Hill, Iqbal, N.E. Karraker, E.S. Kilpatrick, T.A. Langen, J. Langford, K. Lauer, A.J. McCarthy, J. Neale, S. Patel, Patton, C. Southwick, N. Stearrett, N. Steijn, M. Tasleem, J.M. Taylor, and J.R. Vonesh. 2017. Effects of roads and land use on frog distributions across spatial scales and reions in the Eastern and Central United States. Diversity and Distributions.23(2):158-170.
4. Kulmatiski, A., S.R.C. Sprouse^, and H. Beard. 2017. Soil type more than precipitation determines fine-root abundance in savannas of Kruger National Park, South Africa. Plant and Soil. 417(1-2):523-333.

2016

1. Kelsey, K.C., A.J. Leffler, K.H. Beard, J.A. Schmutz, R.T. Choi*, and J.M. Welker. In press. Interactions among vegetation, climate and herbivory control greenhouse gas fluxes in a subarctic coastal wetland. JGR - Biogeosciences.121(12): 2960–2975.
2. Tingley, R., P.J. Mahoney, A.M. Durso, A.G. Taillan, A. Morán-Ordóñez, and K.H. Beard. 2016. Threatened and invasive reptiles are not two sides of the same coin. Global Ecology and Biogeography. 25(9):1050-1060.
3. Costello, M.J., K.H. Beard, R.T. Corlett, G. Cumming, R. Loyola, B. Mea, A.J. Miller-Rushing, R. Pakeman, R.B. Primack. 2016. Field work ethics in biological research: Viewpoint of Biological Conservation editors. Biological Conservation. 203:268-271.
4. Tonini, J.F.R., K.H. Beard, R.B. Ferreira*, and R.A. Pyron. 2016. Fully-sampled phylogenies for squamates reveal evolutionary patterns in extinction risk. Biological Conservation. Special Issue on Reptile Conservation. DOI: 10.1016/j.biocon.2016.03.039
5. Kulmatiski, A., K. H. Beard, J. Grezer, L. Forero, and J. Heavilin. 2016. Plant-soil feedbacks marginally improve predictions of plant biomass in communities. Ecology. 97(8):2064-2073. DOI: 10.1890/15-2037.1
6. Ferreira, R.B.*, K.H. Beard, and M. Crump. 2016. Breeding guilds determine frog response to edge effects in Brazil’s Atlantic Forest. PLOS ONE. 10(12):e0142893.
7. Wallis, A.C.^, R. L. Smith*, and K.H. Beard. 2016. Temporal foraging patterns of non-native Coqui frogs (Eleutherodactylus coqui) in Hawaii. Journal of Herpetology.
8. Ferreira, R.B.*, R. Lourenço-de-Moraes, R.L. Teixeira, and K.H. Beard2016. Frogs associations with bromeliads in a cacao agroecosystem.Northwestern Journal of Zoology. (on-line first): e162501.
9. Mageski, M.M., B. Ferreira*, K.H. Beard, L.C. Costa, P.R. Jesus, C.C. Medeiros, and P.D. Ferreira. 2016. Bromeliad selection of Phyllodytes luteolus (Anura, Hylidae): the influence of plant structure and water quality factors. Journal of Herpetology. 50(1):108-112.DOI: 10.1670/14-166.

2015

1. Mahoney, P.J., K.H. Beard, A.G. Taillian, A.L. Long, A.M. Durso, R.J. Kindermann, N.E. Nolan*, D. Kinka, and H.E. Mohn. 2015. Introduction effort and climate matching outperform species traits as predicators of global establishment success of non-native reptiles. Diversity and Distributions. 21(1): 64-74.
2. Nolan, N.E.*, A. Kulmatiski, H. Beard, and J.M. Norton. 2015. Activated carbon decreases invasive plant growth by mediating plant-microbe. AoB Plants.7:plu072. Special Issue on The Role of Below-Ground Processes in Mediating Plant Invasions.
3. Ferreira, R.B.*, K.H. Beard, R.T. Choi*, and W.C. Pitt. 2015. Diet of the non-native greenhouse frog in Maui. Journal of Herpetology. 49(4):586-593.
4. Ferreira, R.B.*, Faivovich, K.H. Beard, and J.P. Pomal. 2015. The first bromeligenous species of Dendropsophus (Anura: Hylidae) from Brazil’s Atlantic Forest. PLOSONE. DOI: 10.1371/journal.pone.0142893.
5. Warren, C.P.*, A. Kulmatiski, and K.H. Beard. 2015. A combined tracer/evapotranspiration model approach estimates plant water uptake in native and non-native shrub-steppe communities. Journal of Arid Environments. 121(2015):67-78.

2014

1. Kulmatiski, A., A. Anderson-Smith, K.H. Beard, S. Doucette-Riis, M. Mazzacavallo, N. Nolan*, and R. Ramirez. 2014. Most soil trophic guilds increase plant growth: a meta-analytical review. Oikos. 10.1111/oik.01767
2. Cosentino, B.J., D.M. Marsh, K. Jones, J.J. Apodaca, C. Bates, J. Beach, K.H. Beard, K. Becklin, J.M. Bell, C. Crockett, G. Fawson*, J. Fjelsted, K.S. Genet, E.A. Forys, M. Grover, J. Holmes, K. Indeck, N.E. Karraker, E. Kilpatrick, T.A. Langen, S. Mugel, A. Molina, J.R. Vonesh, R. Weaver and A. Willey*. 2014. Roads negatively affect frog and toad distributions throughout the eastern and central United States. Biological Conservation.180:31-38.
3. Kalnicky, E.A.*, M. Brunson, K.H. Beard.  2014.  A social-ecological systems approach to non-native species: Habituation and its impact on management of coqui frogs in Hawaii. Biological Conservation. 180:187-195.
4. Olson, C.A.*, A. Diesmos, and H. Beard. 2014. Geographical Distribution: Eleutherodactylus planirostris (greenhouse frog). Herpetological Review. 45 (4): 652-653.

2013

1. Kulmatiski, A. and K.H. Beard. 2013. Increasing precipitation intensity facilitates woody plant encroachment. Nature Climate Change. 3: 833-837.
2. Hoopes, M., D. Marsh, K.H. Beard, N. Goldberg, A. Aparicio, A. Arbuthnot, B. Hixon, D. Laflower, L. Lee, A. Little, E. Mooney, A Pallette, A. Ravenscraft, S. Scheele, K. Stowe, C. Skyes, R. Watson, B. Yang. 2013. Invasive plants in U.S. National Wildlife Refuges: A coordinated research project using undergraduate ecology students. Bioscience. 63(8): 644-656.
3. Beard, K.H. 2013. Book review of Richard Hobbs et al. editors. 2013. Novel Ecosystems: Intervening in the New World Order. Wiley-Blackwell, Oxford. 380 p. Biological Conservation. 169:157. 
4. Kalnicky, E.A.*, K.H. Beard, and M. Brunson. 2013. Community-level response to habitat structure manipulations: an experimental case study in a tropical ecosystem. Forest Ecology and Management. 307: 313-321.
5. Beard, K.H., C.A. Faulhaber*, F.P. Howe, and T.C. Edwards. 2013. Rodent-mediated interactions among seed species of differing quality in a shrubsteppe ecosystem. Western North American Naturalist. 73(4):426-441.
6. Poessel, S.A., K.H. Beard, C.M. Callahan, R.B. Ferreira*, and E.T. Stevenson. 2013. Biotic acceptance in introduced amphibians and reptiles in Europe and North America. Global Ecology and Biogeography. 22:192-201.
7. Kulmatiski, A. and K.H. Beard. 2013. Root niche partitioning among grasses, saplings, and trees measured using a tracer technique. Oecologia. 171(1): 25-37.

2012

1. O'Neill, E.M.*, K.H. Beard, and M.E. Pfrender. 2012. Cast adrift on an island: introduced populations experience an altered balance between selection and drift. Biology Letters. 8(5):890-893.
2. Kulmatiski, A., K.H. Beard, and J. Heavilin. 2012. Plant-soil feedbacks provide an alternative explanation for diversity-productivity relationships. Proceedings of the Royal Society B. 279: 3020–3026. doi: 10.1098/rspb.2012.0285
3. Ferreira, R.B.*, K.H. Beard†, S.L. Peterson, S.A. Poessel, and C.M. Callahan. 2012. Establishment of introduced reptiles increases with the presence and richness of native congeners. Amphibia-Reptilia. 33 (2012):387-392.
4. Ferreira, R.B.*, C.M. Callahan, S.A. Poessel, and K.H. Beard†. 2012. Global assessment of establishment success for amphibian and reptile invaders. Wildlife Research. 39(7): 637-640.
5. Pitt, W.C., K.H. Beard, and R.E. Doratt. 2012. Management of invasive coqui frog populations in Hawaii. Outlooks on Pest Management. 23(4): 166-169.
6. Crowl, T.A., N. Brokaw, R. Waide, G., Gonzalez, K.H. Beard, E. Greathouse, A.E. Lugo, A.P Covich, J. Lodge, C. Pringle, J. Thompson, G.E. Belovsky. 2012. Chapter 6: When and where biota matter. In: A Caribbean Forest Tapestry: The Multidimensional Nature of Disturbance and Response (N. Brokaw, T. Crowl, A. Lugo, W.M. McDowell, F. Scatena, R.E. Waide, M.R. Willig, eds.) Oxford University Press, Oxford. pp. 272-304.
7. Scatena, F.N., J.F. Blanco, K.H. Beard, R. Waide, A.E. Lugo, N. Brokaw, W. Silver, B. Haines, J. Zimmerman. 2012. Chapter 4: Disturbance regime. In: A Caribbean Forest Tapestry: The Multidimensional Nature of Disturbance and Response (N. Brokaw, T. Crowl, A. Lugo, W.M. McDowell, F. Scatena, R.E. Waide, M.R .Willig eds.) Oxford University Press, Oxford. pp. 164-200.
8. Beard, K.H. and W.C. Pitt. 2012. Chapter 26: Caribbean tree frog (Eleutherodactylus coqui). Pp. 311-319. In: Handbook of Global Freshwater Invasive Species. Earthscan, London. Ed. Robert A. Francis.
9. Olson, C.A.*, K.H. Beard†, and W.C. Pitt. 2012. Pacific Island invasive species: 8. Eleutherodactylus planirostris, the Greenhouse frog (Anura: Leptodactylidae). Pacific Science. 66(3):255-270.
10. Bisrat, S.A., M.A. White, K.H. Beard†, and D.R. Cutler. 2012. Predicting the distribution potential of an invasive Puerto Rican frog (Eleutherodactylus coqui) in Hawaii using remote sensing data. Diversity and Distributions. 18(7):648-660.
11. Choi, R.T.*, and K.H. Beard†. 2012. Coqui frog invasions change invertebrate communities in Hawaii. Biological Invasions. 14(5):939-948.
12. Beard, K.H. and A. Kulmatiski. 2012. Introduction, establishment, and spread: 50 years of invasion ecology since Elton. Book review of David M. Richardson, editor. 2011. Fifty years of invasion ecology: the legacy of Charles Elton. Wiley-Blackwell, Oxford. 432 p. Ecology 93(2): 437-438.
13. Olson, C.A.*, K.H. Beard†, D.N. Koons, and W.C. Pitt. 2012. Detection probability of two introduced frogs in Hawaii: implications for assessing non-native species distributions. Biological Invasions. 14(4): 889-900.
14. Olson, C.A.* and K.H. Beard†. 2012. Diet of the invasive greenhouse frog in Hawaii. Copeia. 1:121-129.

2011

1. O’Neill, E.M.* and Beard, K.H.† 2011. Clinal variation in calls of native and introduced populations of Eleutherodactylus coqui. Copeia 2011(1):18–28.
2. Kulmatiski, A. and K.H. Beard. 2011. Long-term plant growth legacies overwhelm short-term plant growth effects on soil microbial community structure. Soil Biology and Biochemistry 43(3):823-830.
3. Kulmatiski, A., J. Heavilin, and K.H. Beard. 2011. Testing predictions of a three-species plant-soil feedback model. Journal of Ecology 99: 542–550. Rated by F1000. Recommended reading: http://www.journalofecology.org/view/0/index.html

2010

1. Gilbert-Norton, L., R. Wilson, J.M. Stevens, and K.H. Beard†. 2010. Corridors increase movement: A meta-analytical review. Conservation Biology 24(3): 660-668. (†contact author)
2. Kulmatiski, A., K.H. Beard, L.A. Myerson, J.C. Gibson, and K.E. Mock. 2010. Non-native Phragmites australis invasion into Utah wetlands. Western North American Naturalist 70(4): 541–552.
3. Kulmatiski, A., K.H. Beard, R.J.T. Verweij, and E.C. February. 2010. A depth-controlled tracer technique quantifies the location, extent, and timing of water uptake in a sub-tropical savanna. New Phytologist.188(1):199-209.
4. O’Neill, E.M.* and K.H. Beard. 2010. Genetic basis of a color pattern polymorphism in the frog Eleutherodactylus coqui. Journal of Heredity 101(6):703-9.

2009

1. Beard, K.H., E.A. Price* (now Kalnicky), and W.C. Pitt. 2009. Biology and impacts of Pacific Island invasive species: Eleutherodactylus coqui, the Coqui frog (Anura: Leptodactylidae). Pacific Science 63(3): 297–316. (Invited Submission)
2. Tuttle, N.C.*, K.H. Beard†, and W.C. Pitt. 2009. Invasive litter, not an invasive insectivore, determines invertebrate communities in Hawaiian forests. Biological Invasions 11(4):845–855.
3. Peacock, M.M., K.H. Beard†, E.M. O’Neill*, V. Kirchoff, and M.B. Peters. 2009. Strong founder effects and low genetic diversity in introduced populations of Coqui frogs. Molecular Ecology 18: 3603-3615.

2008

1. Bolger, D.T., K.H. Beard, A. Suarez, and T. Case. 2008. Increased abundance of native and non-native spiders with habitat fragmentation. Diversity and Distributions 14(4): 655-665.
2. Beard, K.H., R. Al-Chokhachy, N.C. Tuttle*, and E.M. O’Neill*. 2008. Population density estimates and growth rates of Eleutherodactylus coqui in Hawaii. Journal of Herpetology. 42(4):626-636.
3. Sin, H.*, K.H. Beard†, and W.C. Pitt. 2008. An invasive frog, Eleutherodactylus coqui, increases new leaf production and leaf litter decomposition rates through nutrient cycling in Hawaii. Biological Invasions. 10(3):335-345.
4. Tuttle, N.C.*, K.H. Beard†, and R. Al-Chokhachy. 2008. Aerially applied citric acid reduces an invasive frog. Wildlife Research. 35(7):676-683. (†contact author)
5. Kulmatiski, A., K.H. Beard, J.M. Stevens, and S.M. Cobbold. 2008. Plant-soil feedbacks: A meta-analytical review. Ecology Letters. 11(9): 980-992.
6. Kyle, G.P.*, A. Kulmatiski, and K.H. Beard†. 2008. Influence of pocket gophers mounds on nonnative plant establishment in a shrubsteppe ecosystem. Western North American Naturalist. 68(3):374-381.
7. Kulmatiski, A. and K.H. Beard. 2008. Decoupling plant-growth from land-use legacies in soil microbial communities. Soil Biology and Biochemistry. 40(5):1059-1068.
8. Peters, M., K.H. Beard, C. Hagen, E.M. O’Neill*, K.E. Mock, W.C. Pitt, and T.C. Glenn. 2008. Isolation of microsatellite loci from the coqui frog, Eleutherodactylus coqui. Molecular Ecology. Resources 8:139-141.

2007

1. Beard, K.H. 2007. Diet of the invasive frog, Eleutherodactylus coqui, in Hawaii. Copeia 2007(2): 281-291.
2. Cutler, D.R., T.C. Edwards, K.H. Beard, A. Cutler, K.T Hess, J.C. Gibson, and J.J. Lawler. 2007. Random forests for classification in ecology. Ecology. 88(11):2783-2792.
3. Kyle, G.P.*, K.H. Beard†, and A. Kulmatiski. 2007. Reduced soil compaction enhances establishment of non-native plant species. Plant Ecology. 193:223-232.
4. Rexroad, E.A*, K.H. Beard†, and A. Kulmatiski. 2007. Vegetation responses to 35 and 55 years of native ungulate grazing in shrub-steppe communities. Western North American Naturalist. 67(1):16-25.
5. Velo-Antón, G., P.A. Burrowes, R. Joglar, I. Martínez-Solano, K.H. Beard, and G. Parra-Olea. 2007. Phylogenetic study of Eleutherodactylus coqui (Anura: Leptodactylidae) reveals deep genetic fragmentation in Puerto Rico and origins in Hawaii. Molecular Phylogenetics and Evolution 45(2007):716-728.

2006

1. Kulmatiski, A. and K.H. Beard. 2006. Activated carbon as a restoration tool: Potential for control of invasive plants in abandoned agricultural fields. Restoration Ecology 14(2): 251-257.
2. Beard, K.H. and W.C. Pitt. 2006. Potential predators of an invasive frog (Eleutherodactylus coqui) in Hawaiian forests. Journal of Tropical Ecology 22(4): 345-347.
3. Kulmatiski, A., K.H. Beard, and J.M. Stark. 2006. Exotic communities shift water-use timing in a shrub-steppe ecosystem. Plant and Soil 288(1-2):271-284.
4. Beard, K.H. 2006. Case Study Box: Puerto Rico and Hawaii: Wet tropical forests and the dilemma of coqui frog conservation and eradication. Pp: 135-137. In: Forests and Society: Sustainability and life cycles of forests in human landscapes Eds. K.A. Vogt, J. Honea, D.J. Vogt, M. Andreu, R. Edmonds, J. Berry, R. Sigurdardóttir, T. Patel-Weynand.
5. Kulmatiski, A., K.H. Beard, and J.M. Stark. 2006. Soil history as a primary control on plant invasion in abandoned agricultural fields. Journal of Applied Ecology 43(5):868-876.

2005

1. Beard, K.H. and E.M. O’Neill*. 2005. Infection of an invasive frog Eleutherodactylus coqui by the chytrid fungus Batrachochytrium dendrobatidis in Hawaii. Biological Conservation 126(4): 591-595.
2. Beard, K.H. and W.C. Pitt. 2005. Potential consequences of the coqui frog invasion in Hawaii. Diversity and Distributions 11(5):427-433.
3. Beard, K.H., K.A. Vogt, D.J. Vogt, F.N. Scatena, A. Covich, T.C. Siccama, R. Sigurdardottir, and T.A. Crowl. 2005. Structural and functional responses of a subtropical forest to 10 years of hurricanes and droughts. Ecological Monographs 75(3):345-361.
4. Beard, K.H., Wang, D., Waite, C.E. Decker, K.L.M., Hawley, G.J, DeHayes, D.D., Hughes, J. W., Cumming, J.R. 2005. Quantifying the importance of abiotic and biotic factors in controlling nutrient export from a developing ecosystem. Ecosystems 8:210-224.

2004

1. Kulmatiski, A., K.H. Beard, and J.N. Stark. 2004. Soil-mediated control on weed establishment and growth. Weed Technology 18: 1353-1358 Suppl. S.
2. Kulmatiski, A. and K.H. Beard. 2004. Reducing sampler error in soil research. Soil Biology and Biochemistry 36(2): 383-385.

2003

1. Beard, K.H., A.K. Eschtruth, K.A. Vogt, D.J. Vogt, and F.N. Scatena. 2003. The effects of the frog Eleutherodactylus coqui on invertebrates and ecosystem processes at two scales in the Luquillo Experimental Forest, Puerto Rico. Journal of Tropical Ecology (37)19: 607-617.
2. Beard, K.H., S. McCullough, and A.K. Eschtruth. 2003. Quantitative assessment of habitat preferences for the Puerto Rican terrestrial frog, Eleutherodactylus coqui. Journal of Herpetology 37(1):10-17.
3. Kulmatiski, A., D.J. Vogt, T.G. Siccama, and K.H. Beard. 2003. Detecting nutrient pool changes in rocky forest soils. Soil Science Society of America Journal 67:1282-1286.

Before 2003

1. Beard, K.H., K.A. Vogt, and A. Kulmatiski. 2002. Top-down effects of a terrestrial frog on nutrient concentrations in a subtropical forest Oecologia 133: 583-593.
2. Vogt, K. A., K.H. Beard, S. Hammann, J. L. O'Hara, D. J. Vogt, F.N. Scatena, and B. A. Parry. 2002. Indigenous knowledge informing management of tropical forests: the link between rhythms in plant secondary chemistry and lunar cycles. Ambio 35: 485-490.
3. Beard, K.H. 2001. The effect of a juvenile terrestrial frog, Eleutherodactylus coqui, on the decomposer food web and leaf litter decomposition rate in the wet forest of Puerto Rico.TRI News 20:18-21.
4. Vogt, K. A., O. Schmitz, O., K.H. Beard, J. L. O’Hara, and M. Booth. 2001. Conservation Biology, Contemporary. pp. 865-881. In: Encyclopedia of Biodiversity. Vol.1 (S. Levin, Ed.) Academic Press.
5. Skelly, D.K., K.H. Beard and N. J. Hengarten. 2000. Animal-distribution modeling in gap analysis: an evolving science. Conservation Biology 14(5):1224-1225.
6. Beard, K.H., N. Hengarten and D. K. Skelly. 1999. Effectiveness of predicting breeding bird distributions using probabilistic models. Conservation Biology 13(5):1108-1116.
7. Kiesecker, J. M., D. K Skelly, K.H. Beard and E. Presser. 1999. Behavioral reduction of infection risk. Proceedings of the National Academy of Sciences 96(16):9165-9168.
8. Beard, K.H. and P. T. DePriest. 1996. Genetic variation within and among mats of the reindeer-lichen, Cladina subtenuis (des abb.) Hale & W.Culb. Lichenologist 28(2):171-182.